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mx id: 3937 | OBO id: HAO:0000395 | URI: http://purl.obolibrary.org/obo/HAO_0000395
harpe synonyms: distal segment of gonopod, dististyle, gonoharpe, gonosquama, gonostylus, harpago, harpes, harpide, palette, paramere, squama, telomere
Defined (both definition and relationships must be met)
Definition:
The sclerite that is located distally on the gonostyle/volsella complex and does not connected to the cupula and to the volsella by conjunctiva or muscles.
written by: Mikó, I., M. J. Yoder, E. Johannes, and A. R. Deans. 2012. Male terminalia of Ceraphronoidea: diversity in a monotonous system. Insect Systematics and Evolution in prep.

Relationships / properties:
The harpe is a sclerite.
The harpe is part of the cuticle. The harpe is part of the external male genitalia.

Figures
Gasteruptiidae
Trigonalidae ; Orthogonalys pulchella Cresson, 1867; Orthogonalys pulchella [from: 11]
Masner lubomirus

Label usage (sensu)
harpe by Crampton, G. C. 1919. The genitalia and terminal abdominal structures of males, and the terminal structures of the larvae of ‘chalastogastrous’ Hymenoptera. Proceedings of the Entomological Society of Washington 21:129-151.
paramere by Snodgrass, R. E. 1941. The male genitalia of Hymenoptera. Smithsonian Miscellaneous Collections 99:1-86.
palette by Boulangé, H. 1924. Recherches sur l’appareil copulateur des Hyménoptères et spécialement des Chalastogastres. Mémoires et Travaux de la Faculté Catholique de Lille 28:1-444.
gonosquama by Peck, O. 1937. The male genitalia in the Hymenoptera (Insecta), especially the family Ichneumonidae. Canadian Journal of Research, Section D, Zoological Sciences 15:221-274.
telomere by Snodgrass, R. E. 1957. A revised interpretation of the external reproductive organs of male insects. Smithsonian Miscellaneous Collections 135:1-60.
harpide by Birket-Smith, S. J. R. 1981. The male genitalia of Hymenoptera – a review based on morphology in Dorylidae (Formicoidea) . Entomologica Scandinavica, Supplement 15:377-397.
distal segment of gonopod by Crampton, G. C. 1919. The genitalia and terminal abdominal structures of males, and the terminal structures of the larvae of ‘chalastogastrous’ Hymenoptera. Proceedings of the Entomological Society of Washington 21:129-151.
dististyle by Bohart, R. M., and A. S. Menke. 1976. Sphecid Wasps of the World, A Generic Revision. University of California Press, Berkeley, CA 695 pp.
harpide by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
harpes by Ross, H. H. 1945. Sawfly genitalia: terminology and study techniques. Entomological News (Philadelphia) 56:261-265.
gonostylus by Smith, E. L. 1970. Evolutionary morphology of external insect genitalia 2 Hymenoptera. Annals of the Entomological Society of America 63:1-27.
gonostylus by Michener, C. D. 1965. A classification of the bees of the Australian and South Pacific regions. Bulletin of the American Museum of Natural History 130:1-362.
harpe by Ross, H. H. 1937. A generic classification of the nearctic sawflies (Hymenoptera: Symphyta). Illinois Biological Monographs 15:1-173.
harpes by Königsmann, E. 1977. Das phylogenetische System der Hymenoptera Teil 2: Symphyta. Deutsche Entomologische Zeitschrift 24:1-40.
harpe by Schulmeister, S. 2001. Male genitalia and copulation in Hymenoptera. Acta Zoologica (Stockholm) 82:331-349.
paramere by Königsmann, E. 1976. Das phylogenetische System der Hymenoptera Teil 1:Einführung, Grundplanmerkmale, Schwestergruppe und Fossilfunde. Deutsche Entomologische Zeitschrift 23:253-279.
paramere by Schulmeister, S. 2001. Male genitalia and copulation in Hymenoptera. Acta Zoologica (Stockholm) 82:331-349.
palette by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
gonosquama by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
gonostylus by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
telomere by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
harpes by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
distal segment of gonopod by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
dististyle by Curators, H. A. O. 2009. The Hymenoptera Anatomy Ontology Curation Team. Hymenoptera Anatomy Ontology.
harpe by Miko, I. 2009. -2014 Curator. Hymenoptera Anatomy Ontology.
squama by Townes, H. 1957. Nearctic wasps of the subfamilies Pepsinae and Ceropalinae. Bulletin of the United States National Museum 209:1-286.
gonoharpe by Ohl, M., C. Rasmussen, and I. Mikó. 2010. HAO working group..
harpago by Yoshimura, M., and B. Fisher. 2011. A revision of male ants of the Malagasy region (Hymenoptera: Formicidae): Key to genera of the subfamily Dolichoderinae. Zootaxa 2794:1-34.
harpago by Snodgrass, R. E. 1957. A revised interpretation of the external reproductive organs of male insects. Smithsonian Miscellaneous Collections 135:1-60.
Comments and tags
comment added over 6 years ago by Istvan Miko
Harpago has been used for a possibly nonhomologous and nonequivalent structure in Glossary of mosquito anatomy.
  Reference: None provided.
  Cross reference: None provided.

comment added over 7 years ago by Matt Yoder
Harpe is the apicalmost sclerite of the male genitalia in Lepidoptera.
  Reference: Crampton, G. C. 1919. The genitalia and terminal abdominal structures of males, and the terminal structures of the larvae of ‘chalastogastrous’ Hymenoptera. Proceedings of the Entomological Society of Washington 21:129-151.
  Cross reference: None provided.

alternative definition added over 4 years ago by Istvan Miko
The sclerite that is connected distally to the gonostipes.
  Reference: Miko, I. 2009. -2014 Curator. Hymenoptera Anatomy Ontology.
  Cross reference: None provided.


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